Longtooth grouper (Epinephelus bruneus)

General data

Main name: Longtooth grouper
Local names: Mud grouper, Kelp grouper, Jabari, Kue
Climates: Tropical, Subtropical
Habitat: Saltwater
Native: Asia
Distribution: Pacific Ocean, South China Sea, East China Sea, Sea of Japan (East Sea)

Classification

Description

Epinephelus bruneus, commonly known as the longtooth grouper, kelp grouper or jabari is a species of marine ray-finned fish, a grouper from the subfamily Epinephelinae which is part of the family Serranidae, which also includes the anthias and sea basses.

It is found in northwest Pacific in eastern Asia, where the species is known by the traditional name "Kue".

Longtooth grouper has an elongate body which has a standard length which is 3.0 to 3.6 times its depth. The dorsal profile of the head between the eyes is convex. The preopercle has an angle where the serrations are notably enlarged. There is a small spine on the upper edge of the gill cover and this upper edge is convex.

The dorsal fin contains 11 spines and 13-15 soft rays while the anal fin has 3 spines and 8 soft rays.

It has 64-72 scales in its lateral line.

The larger adults are dark greyish brown in colour, and may be marked with faint blotches on their backs, although these may be absent. They also have a covering of small grey spots on their bodies which form short longitudinal lines and creating a mottled pattern.

The lower margin of the anal fin and the lower corner of caudal fin have white edges.

The juveniles are pale yellowish brown in colour, with 6 irregular, diagonal dark bars within which there are irregular pale spots. The first of these bars extends from nape to eye and the last is on the caudal peduncle. There are 3 dark brown bands which radiate from lower part of eye and some juveniles have greenish yellow membranes between the rearmost spines of the dorsal fin.

The maximum published total length for this species is 136 centimetres (54 in), although the most common length is around 60 centimetres (24 in), and the maximum recorded weight is 33 kilograms (73 lb).

Epinephelus bruneus is found in the Western Pacific Ocean where it occurs from southern Japan and Taiwan and along the coast of Asia from southern Korea to Hainan and Hong Kong in southern China.

Taxonomy of Fishes

Osteichthyes - Bony fishes->
- Actinistia - Lobe-finned fishes->
- Actinopteri - Ray-finned fishes->
  - Perciformes - Perches->
    
    Percidae - Perches->
    
    Pomatomidae - Bluefishes->
    
    Serranidae - Sea basses->
    
    Pomacentridae - Damsel and clownfishes->
    
    Latidae - Lates perches->
    
    Centropomidae - Snooks->
    
    Lutjanidae - Snappers->
    
    Haemulidae - Grunts->
    
    Artedidraconidae - Barbled plunderfishes->
    
    Embiotocidae - Surfperches->
    
    Glaucosomatidae - Pearl perches->
    
    Malacanthidae - Tilefishes->
    
    Polynemidae - Threadfin->
    
    Gerreidae - Mojarras->
    
    Priacanthidae - Bigeyes->
    
    Monodactylidae - Moonyfishes->
    
    Centrogenyidae - False scorpionfishes->
    
    Epinephelidae - Groupers->
    
    Epinephelus - Cloudy groupers->
    
    Atlantic goliath grouper (Epinephelus itajara)
    
    Dusky grouper (Epinephelus marginatus)
    
    Goldblotch grouper (Epinephelus costae)
    
    Red grouper (Epinephelus morio)
    
    Nassau grouper (Epinephelus striatus)
    
    Malabar grouper (Epinephelus malabaricus)
    
    Maori rockcod (Epinephelus undulatostriatus)
    
    Blacktip grouper (Epinephelus fasciatus)
    
    One-blotch grouper (Epinephelus melanostigma)
    
    Comet grouper (Epinephelus morrhua)
    
    Pacific goliath grouper (Epinephelus quinquefasciatus)
    
    Giant grouper (Epinephelus lanceolatus)
    
    Honeycomb grouper (Epinephelus merra)
    
    Potato grouper (Epinephelus tukula)
    
    Blue-and-yellow grouper (Epinephelus flavocaeruleus)
    
    White grouper (Epinephelus aeneus)
    
    Epaulet grouper (Epinephelus stoliczkae)
    
    Rock hind (Epinephelus adscensionis)
    
    Hong Kong grouper (Epinephelus akaara)
    
    White-edged grouper (Epinephelus albomarginatus)
    
    Banded grouper (Epinephelus amblycephalus)
    
    Spotted grouper (Epinephelus analogus)
    
    Catface grouper (Epinephelus andersoni)
    
    Areolate grouper (Epinephelus areolatus)
    
    Yellow grouper (Epinephelus awoara)
    
    Frostback rockcod (Epinephelus bilobatus)
    
    Duskytail grouper (Epinephelus bleekeri)
    
    Palemargin grouper (Epinephelus bontoides)
    
    Longtooth grouper (Epinephelus bruneus)
    
    Dogtooth grouper (Epinephelus caninus)
    
    Moustache grouper (Epinephelus chabaudi)
    
    Tonga grouper (Epinephelus chlorocephalus)
    
    Brownspotted grouper (Epinephelus chlorostigma)
    
    Epinephelus clippertonensis (Epinephelus clippertonensis)
    
    Whitespotted grouper (Epinephelus coeruleopunctatus)
    
    Orange-spotted grouper (Epinephelus coioides)
    
    Coral grouper (Epinephelus corallicola)
    
    Brokenbar Grouper (Epinephelus craigi)
    
    Speckled blue grouper (Epinephelus cyanopodus)
    
    Saddletail grouper (Epinephelus daemelii)
    
    Spinycheek grouper (Epinephelus diacanthus)
    
    Speckled hind (Epinephelus drummondhayi)
    
    Dotted grouper (Epinephelus epistictus)
    
    Cloudy grouper (Epinephelus erythrurus)
    
    Rock grouper (Epinephelus fasciatomaculosus)
    
    Barred-chest grouper (Epinephelus faveatus)
    
    Brown-marbled grouper (Epinephelus fuscoguttatus)
    
    Darkmargin grouper (Epinephelus fuscomarginatus)
    
    Multispotted grouper (Epinephelus gabriellae)
    
    Epinephelus geoffroyi (Epinephelus geoffroyi)
    
    Dungat grouper (Epinephelus goreensis)
    
    Red hind (Epinephelus guttatus)
    
    Bridled grouper (Epinephelus heniochus)
    
    Starspotted grouper (Epinephelus hexagonatus)
    
    Blacksaddle grouper (Epinephelus howlandi)
    
    Somali grouper (Epinephelus indistinctus)
    
    Pacifc insular grouper; (Epinephelus insularis)
    
    Marquesan grouper (Epinephelus irroratus)
    
    Brownspotted grouper (Epinephelus japonicus)
    
    Fivebar grouper (Epinephelus kupangensis)
    
    Starry grouper (Epinephelus labriformis)
    
    Striped grouper (Epinephelus latifasciatus)
    
    Mystery grouper (Epinephelus lebretonianus)
    
    Longspine grouper (Epinephelus longispinis)
    
    Snubnose grouper (Epinephelus macrospilos)
    
    Highfin grouper (Epinephelus maculatus)
    
    Speckled grouper (Epinephelus magniscuttis)
    
    Netfin grouper (Epinephelus miliaris)
    
    White-blotched grouper (Epinephelus multinotatus)
    
    White-streaked grouper (Epinephelus ongus)
    
    Dot-dash grouper (Epinephelus poecilonotus)
    
    Smallscaled grouper (Epinephelus polylepis)
    
    Camouflage grouper (Epinephelus polyphekadion)
    
    White-dotted grouper (Epinephelus polystigma)
    
    Striped-fin grouper (Epinephelus posteli)
    
    Longfin grouper (Epinephelus quoyanus)
    
    Oblique-banded grouper (Epinephelus radiatus)
    
    Red-tipped grouper (Epinephelus retouti)
    
    Halfmoon grouper (Epinephelus rivulatus)
    
    Sixbar grouper (Epinephelus sexfasciatus)
    
    Surge grouper (Epinephelus socialis)
    
    Foursaddle grouper (Epinephelus spilotoceps)
    
    Black-dotted grouper (Epinephelus stictus)
    
    Seamount grouper (Epinephelus suborbitalis)
    
    Summan grouper (Epinephelus summana)
    
    Greasy grouper (Epinephelus tauvina)
    
    Yellowspotted grouper (Epinephelus timorensis)
    
    Threespot grouper (Epinephelus trimaculatus)
    
    Plump grouper (Epinephelus trophis)
    
    Reticulate grouper (Epinephelus tuamotuensis)
    
    Wavy-lined grouper (Epinephelus undulosus)
    
    Mycteroperca - Groupers ->
    
    Cephalopholis - Scaleheaded groupers->
    
    Hyporthodus - Hyporthodus->
    
    Alphestes - Alphestes->
    
    Variola - Variola->
    
    Anyperodon - Anyperodon->
    
    Gonioplectrus - Gonioplectrus->
    
    Plectropomus - Coral groupers->
    
    Gracila - Gracila->
    
    Aethaloperca - Aethaloperca->
    
    Chromileptes - Humpback groupers->
    
    Dermatolepis - Dermatolepis->
    
    Saloptia - Saloptia->
    
    Triso - Triso->
    
    Cebidichthyidae - Cebidichthyidae->
    
    Anthiadidae - Anthiadidae->
    
    Ambassidae - Asiatic glassfishes->
    
    Bathydraconidae - Antarctic dragonfishes->
    
    Bembropidae - Duckbill flatheads->
    
    Bovichtidae - Thornfishes->
    
    Caesionidae - Fusiliers->
    
    Caesioscorpididae - Blowhole perches->
    
    Centracanthidae - Picarels->
    
    Cepolidae - Bandfishes->
    
    Channichthyidae - Crocodile icefishes->
    
    Congrogadidae - Eelblennies->
    
    Datnioididae - Freshwater tripletails->
    
    Dinopercidae - Cavebasses->
    
    Eleginopidae - Patagonian blennies->
    
    Grammatidae - Basslets->
    
    Grammistidae - Soapfishes->
    
    Hapalogenyidae - Barbeled grunters->
    
    Harpagiferidae - Spiny plunderfishes->
    
    Lactariidae - False trevallies->
    
    Latilidae - Deepwater tilefishes->
    
    Leptobramidae - Beachsalmon->
    
    Liopropomatidae - Painted basslets->
    
    Niphonidae - Ara groupers->
    
    Nototheniidae - Cod icefishes->
    
    Opistognathidae - Jawfishes->
    
    Oxylebiinae - Painted greenlings->
    
    Plesiopidae - Roundheads->
    
    Pseudaphritidae - Catadromous icefishes->
    
    Pseudochromidae - Dottybacks->
    
    Ptereleotridae - Dartfishes->
    
    Toxotidae - Archerfishes->
  - Cypriniformes - Carps->
  - Esociformes - Pikes->
  - Salmoniformes - Salmons and Trouts->
  - Siluriformes - Catfishes->
  - Scorpaeniformes - Mail-cheeked fishes->
  - Gasterosteiformes - Sticklebacks->
  - Gadiformes - Cods->
  - Anguilliformes - Eels and morays->
  - Clupeiformes - Herrings->
  - Syngnathiformes - Pipefishes and Seahorses->
  - Gobiiformes - Gobies->
  - Mugiliformes - Mullets->
  - Carangiformes - Jacks->
  - Beloniformes - Needlefishes->
  - Labriformes - Wrasses->
  - Scombriformes - Mackerels->
  - Mulliformes - Goatfishes->
  - Pleuronectiformes - Flatfishes->
  - Aulopiformes - Grinners->
  - Blenniiformes - Blennies->
  - Cyprinodontiformes - Toothcarps->
  - Osmeriformes - Smelts->
  - Centrarchiformes - Basses and sunfishes->
  - Cichliformes - Cichlids->
  - Characiformes - Characins->
  - Osteoglossiformes - Bony tongues->
  - Elopiformes - Tarpons and tenpounders->
  - Anabantiformes - Gouramies and snakeheads->
  - Gymnotiformes - South American knifefish->
  - Istiophoriformes - Barracudas->
  - Albuliformes - Bonefishes->
  - Lepidogalaxiiformes - Salamanderfishes->
  - Lophiiformes - Anglerfishes->
  - Spariformes - Breams and porgies->
  - Trachiniformes - Weeverfishes->
  - Moroniformes - Temperate basses->
  - Acanthuriformes - Surgeonfishes->
  - Zeiformes - Dories->
  - Tetraodontiformes - Puffers and filefishes->
  - Holocentriformes - Squirrelfishes->
  - Dactylopteriformes - Flying gurnards->
  - Lampriformes - Lamprids->
  - Galaxiiformes - Southern smelts->
  - Batrachoidiformes - Toadfishes->
  - Hiodontiformes - Mooneyes->
  - Percopsiformes - Trout-perches->
  - Callionymiformes - Dragonets->
  - Atheriniformes - Silversides->
  - Notacanthiformes - Spiny eels->
  - Saccopharyngiformes - Swallowers and Gulpers->
  - Alepocephaliformes - Slickheads and tubeshoulders->
  - Gonorynchiformes - Milkfishes->
  - Argentiniformes - Marine smelts->
  - Stomiiformes - Lightfishes and dragonfishes->
  - Ateleopodiformes - Jellynose fishes->
  - Myctophiformes - Lanternfishes->
  - Stylephoriformes - Tube-eyes->
  - Polymixiiformes - Beardfishes->
  - Beryciformes - Sawbellies->
  - Trachichthyiformes - Rughies->
  - Ophidiiformes - Cusk-eels->
  - Kurtiformes - Nurseryfishes & Cardinalfishes->
  - Synbranchiformes - Swamp eels->
  - Gobiesociformes - Clingfishes->
  - Acropomatiformes - Oceanic basses->
- Cladistii - Bichirs->
- Dipnoi - Lungfishes->
- Chondrostei - Sturgeons and Paddlefishes->
- Holostei - Gars and Bowfins->
Chondrichthyes - Cartilaginous fishes->
Agnatha - Jawless fishes->

Longtooth grouper (Epinephelus bruneus)

General data

Classification

Description

Taxonomy of Fishes

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